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sand
fiddler
crab
calling
wetland
mangrove
species
claw
gaomei
marsh
nature
habitat
swamp
indopacific
beach
animal
crustacean
tidal
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male
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wildlife
malaysia
shore
atlantic
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calico
tubuca
leptuca
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uca
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pugnax
natural
thailand
taichung
wave
indonesia
drumming
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season
display
courtship
ocypodidae
marine
taiwan