Neohelice granulata crab walking sideways over mudflat, slipping into water

crustacea neohelice granulata walking sideways intertidal zone estuary biology euryhalinity burrowing semiterrestrial crab chasmagnathus granulata highly studied species ecological research mangroves crab filter feeding into water salt marsh ecology physiology emergent animal model biochemical research malacostraca decapoda brachyura varunidae crustacean model species in biological fields semiterrestrial habit pincers mudflats decapod crustaceans thorax exoskeleton chitin pair of chelae claws camouflage hide away walk underwater animal creature legs outside outdoors eyestalk walking scientific studies marsh creature large pincers nature