Portrait Of A Male African Bullfrog Inflating His Vocal Sacs In A Puddle

male african bullfrog call mate mating vocal sacs throat expanding inflated pond puddle shallow animal lake habitat season reflection bullfrog african botswana frog outdoor creature environment nature mouth video ecology wet toad adapted southafrica eye secretion natural adspersus behaviour aquatic large secretive vocal pyxicephalus metamorphoses giant slimy moist anura africa breeding