Wild crustacean found in its natural habitat, a fiddler crab foraging and sipping minerals in a puddle of water in the muddy tidal flat at Gaomei wetlands preservation area, Taichung, Taiwan

crab ocypodidae crustacean male sediment puddle water female mud wetland wildlife swamp species shell muddy environment arcuatus marsh animal sand ecosystem mangrove tidal claw gelasimus beach habitat forage arcuata tubuca eat fauna soil marine nature flat coastal ground sexually dimorphic natural gaomei fiddler calling preservation borealis taiwan asymmetric bowed taichung